EU546815	8636	417	1220	URA3
EU546815	8636	2274	2431	palA lagging strand origin
EU546815	8636	2570	3220	cat194
EU546815	8636	3389	3772	rep
EU546815	8636	3427	3555	oriV for pC194
EU546815	8636	3677	4366	pre
EU546815	8636	4613	4699	palB
EU546815	8636	5452	6053	ColE1 high copy variant
EU546815	8636	6234	7094	bla